THE WORLD BIRDS - An Online Bird Book
BUCEROTIFORMES
The order
Bucerotiformes contains the
hornbills,
hoopoe and
woodhoopoes. In addition to DNA comparisons, these three groups of birds have similar skulls and leg bones. However, the hornbills are very different in appearance and the current thought is that they have a different common ancestor from the hoopoes and woodhoopoes.
HORNBILLs
Order Bucerotiformes Family Bucerotidae
Hornbills are a family of bird found in tropical and subtropical Africa and Asia. They are characterized by a long, down-curved bill which is frequently brightly-colored and sometimes has a casque on
the upper mandible. The heavy bill is is used for catching prey, constructing the nest, preening, and also for fighting. Because it is so heavy it is supported by powerful neck muscles and and two neck vertebrae have fused to add support.
A feature unique to the hornbills is the casque, a usually hollow structure that is attached to the top of the bill. In some species it is barely
perceptible and appears to serve no function beyond reinforcing the bill. In other species it is quite large, is reinforced with bone, and has openings between the hollow center serving as a resonator
for calls.
Image by Shantanu Kuveskar
Hornbills show considerable variation in size, from the black dwarf hornbill (Tockus hartlaubi) which is 32 cm long and 100 grams to the southern ground hornbill which has an average weight of 3700 g. Males are always heavier than their female counterparts, though the extent to which this is true varies according to species. Males also have longer bills, casques, and wings.
The family is omnivorous, feeding on fruit, insects, and small animals. Species that eat mainly fruit are generally found in forests as that is where most of the fruit is. One of their favorite fruits is figs. They will also visit cultivated fruit trees that are not located in forests. The hornbills that prefer insects and small animals prefer more open areas where they can sight and thence catch prey. They cannot swallow food caught at the tip of the beak as their tongues are too short to manipulate it, so they toss it back to the throat
with a jerk of the head. Because of their preference for fruit, hornbills play an important role in dispersing their seeds.
Hornbills nest in natural cavities in trees and sometimes cliffs. These cavities are usually natural, often caused by a large limp falling off and subsequent rot resulting in a hole. They are known to use nests that were perviously constructed by woodpeckers and barbets. One of the most amazing habits of the hornbills is that the breeding female will, usually with the help of her mate, partially seal herself into the nest cavity. Hornbills use their bills in this sealing process to construct a wall made out of material such as mud, droppings, and fruit pulp. A space is left in the wall so the male can supply the female and the chicks with whole or regurgitated food. The female will stay in the partially sealed cavity for months while the eggs are laid, and incubated. In some species the female will continue living in the sealed cavity until the chicks fledge, in other species she will leave a few weeks after the first chick hatches and then help her mate feed the young. The female molts while she is sealed in the nest cavity. Think how the nest must smell after months of being almost entirely closed, even containing perhaps a dead chick or two! Why do they go to this extreme behavior? It must be to increase the odds of the chicks surviving attacks from predators.
Male feeding female at nest by JM Garg
Many Asian hornbills are threatened by habitat loss, as they tend to require mature trees that provide natural cavities for nesting. Hunting provides an added stress to the population. The African hornbills are generally maintain a stable population, but some of these species are also in trouble. Exacerbating the hornbills stress from humans, many hornbills only produce a maximum of one surviving offspring every year. The first born chick is usually 3 to 5 days older than its sibling; as result it is considerably larger and usually gets the majority of food so the younger chick dies. In essence, the second chick was an insurance policy. If the first born is defective, then chick number 2 gets lucky, otherwise life is hard.
The New World toucans of family Ramphastidae are the birds that most closely resemble the hornbills. Like the hornbills, the toucans have massive brightly colored bills. But these two heavy-billed families are not related. Rather, convergent evolution led them to develop similar bills. The hornbills closest relatives are the hooe and wood hoopies which are in the same order as the hornbills,
Bucerotiformes.
Genus Aceros - 1 species
Hornbill,_Rufous-necked Aceros nipalensis
Image by: 1)
Ron_Knight 2) Debmayukh285 - India 3)
Kalyanvarma 4)
Ujjal_Ghosh - India
1) Pair
2) Female 3, 4) Male
Genus Anorrhinus
They are found in forests of Southeast Asia (just barely extending into adjacent parts of India and China). They are social and typically seen in groups, but only the dominant pair are believed to breed, while other group members act as helpers. Because of habitat destruction, all three species are listed as Near Threatened.
Hornbill,_Brown Anorrhinus austeni
Image by: 1)
Pkspks 2, 3, 4) LonelyShrimp- Thailand
1) Female 2 - 4) Male
Hornbill,_Bushy-crested Anorrhinus galeritus
Image by: 1)
Dick_Culbert 2)
Tony_Castro 3)
David_Cook - Borneo
Hornbill,_Rusty-cheeked also
Tickell's Hornbill Anorrhinus tickelli
Image by: 1)
Mike 2)
Jason Thompson
Genus Anthracoceros
These are the blac Asian hornbills. They all have mainly black plumage. Some have white bare skin around the eyes and a white throat patch. And some also have a white tail.
Hornbill,_Black Anthracoceros malayanus Found, Asia
Image by: 1)
Mahbob
Yusof 2)
Neil Phillips- the London Zoo 3)
Mike_Prince - Borneo 4)
Petra_Karstedt
1) Female 2, 3, 4) Male
Hornbill,_Malabar_Pied Anthracoceros coronatus
Image by: 1)
Uday_Agashe 2)
Steve Garvie - Sri Lanka
3, 4)
Thimindu Goonatillake - Sri Lanka
1) Pair, male on left 2) Female 3, 4) Male
Hornbill,_Oriental Pied Anthracoceros albirostris
Image by: 1)
LonelyShrimp - Thailand 2)
Mahbob Yusof 3)
Dick Daniels - Bronx Zoo 4)
Paul_Williams - Malaysia
1) Male left, female right 2) Female 3) Male
Hornbill,_Palawan Anthracoceros marchei
Image by: 1)
Limchiu 2)
Raniel_jose_castaneda
1) Male at top
Hornbill,_Sulu Anthracoceros montani
Image by:
1)
Biodiversity_Heritage_Library
Genus Berenicornis - 1 species
Hornbill,_White-crowned Berenicornis comatus
Image by:
1, 2) Choy Wai Mun - Nalaysia
3)
Tapir Girl - Franklin Park Zoo
Bernard_Dupont Peter_Steward
Genus Buceros
These are large hornbills which all have a large, hollow casque. They have the largest wingspan of all the hornbills, up to 1.8 meters.
Hornbill,_Great also
Great Indian Hornbill Buceros bicornis
Image by: 1)
Eric_Kilby 2)
Joachim Muller 3)
Stavenn - Bronx Zoo
4) Lip Kee - Malaysia
1) Female 2) Male
Hornbill,_Helmeted Buceros vigil
Image by: 1)
Farquhar 2)
Doug Janson
Hornbill,_Rhinoceros Buceros rhinoceros
Image by: 1)
Perry Quan - Dallas Zoo 2)
Steve_Wilson 3)
Dick Daniels -
National Aviary 4)
Sandy Cole -
National Aviary
1, 2) Female (white iris) 3, 4) Male
Hornbill,_Rufous Buceros hydrocorax
Image by:
1)
Frank Wouters 2)
Fiorellino 3)
Magalhaes 4)
NCAA_Official Olaf_Oliviero_Riemer
1) Juvenile
2, 3) Female 4, 5) Male
Genus Bycanistes
The African hornbills have black and white plumage. The plumage of the sexes is similar, but the casque of the male is larger than that of the female. They have often been included in the genus
Ceratogymna,
Hornbill,_Black-and-white-casqued Bycanistes subcylindricus
Image by:
1, 3) Nik_Borrow - Uganda 2)
Bernard_Dupont 4)
Dante Alighieri - Bronx Zoo
1, 2) Female 3, 4) Male
Hornbill,_Brown-cheeked Bycanistes cylindricus
Image by:
1)
John Gerrard Keulemans 2)
Dimitry_Naumenko 3)
Hans Olofsson - Uganda 4)
Nik_Borrow - Ghana
Hornbill,_Piping Bycanistes fistulator
Image by: 1
) Dick Daniels -
World of Birds
, South Africa 2)
Moni Sertel 3)
Dotun55 - Nigeria
Hornbill,_Silvery-cheeked Bycanistes brevis
Image by: 1)
Katy Silberger 2, 3) Dick Daniels -
Sylvan Heights 4)
Nik_Borrow - Tanzania
1) Female 2 - 4) Male
Hornbill,_Trumpeter Bycanistes bucinator
Image by: 1)
Dick Daniels -
World of Birds, South Africa 2)
Eric_Kilby 3)
Ian White 4)
Lip Kee - Botswana
1) Female 2, 3, 4) Male
Hornbill,_White-thighed Bycanistes albotibialis
Image by:
1)
John Gerrard Keulemans 2)
Sergey_Pisarevskiy - Uganda 3)
Nik_Borrow - Uganda
3) Pair
Genus Ceratogymna
Hornbill,_Black-casqued Ceratogymna atrata
Image by: 1)
Daderot 2)
Nik_Borrow - Ghana 3)
Josh_More
1) Pair 2) Female
3) Male
Hornbill,_Yellow-casqued Ceratogymna elata
Image by:
1)
Denise Chan 2)
Josh_More
1) Female
2) Male
Genus Horizocerus
Hornbill,_Black_Dwarf Horizocerus hartlaubi
Image by: 1)
Daniel Giraud Elliot 2)
Nik_Borrow - Ghana
Hornbill, White-crested also
Horizocerus albocristatus
Image by: 1)
Sandy Cole - San Diego Zoo 2,
4) Dick Daniels - Bronx Zoo, San Diego Zoo
3) Charlie Westerinen
Genus Lophoceros
These hornbills are all native to Africa. They are small hornbills, about 50 cm in length.
Hornbill,_African_Pied Lophoceros fasciatus
Image by: 1)
Nik Borrow - Guinea
2, 4) Matti_Suopajärvi - Gambia 3)
Anneli
Moeller
Hornbill,_Bradfield's Lophoceros bradfieldi
Image by: 1)
Ron_Knight 2)
Charlie Westerinen - Hwange National Park in Zimbabwe 3) I
an White - Botswana
Hornbill,_Crowned Lophoceros alboterminatus
Image by: 1)
Bernard_Dupont - South Africa 2)
Derek Keats 3) Sergey_Yeliseev 4)
Nik_Borrow -
Uganda
Hornbill,_African grey Lophoceros nasutus
Image by: 1)
Dominic_Sherony 2, 3) Charles_J_Sharp - Namibia, Gambia 4) Nik_Borrow
1, 2) Female 3, 4) Male
Hornbill,_Hemprich's Lophoceros hemprichii
Image by: 1)
Steve Evans Ethopia 2)
Alastair Rae 3)
Michael and Helen Cox - Kenya
Hornbill,_Pale-billed Lophoceros pallidirostris
Image by:
1)
Peter_Steward - Malawi
2, 3, 4) Nik_Borrow - Malawi, Malawi, Tanzania
Genus Ocyceros
These relatively small hornbills are found either in India or the neiighboring island of Sri Lanka. They have grey plumage and either small or nonexistant casques.
Hornbill,_Indian_grey Ocyceros birostris
Image by:
1) J M Garg -
Hyderabad, India 2)
Raju_Kasambe 3)
Tarique_Sani 4)
photosinframes
2) Male supplying food to nest inmates
Hornbill,_Malabar_grey Ocyceros griseus
Image by:
1, 3) Lip Key 2) Rathika Ramasamy 4)
Nagesh_Kamath
1) Female 2, 3) Male
Hornbill,_Sri_Lanka_grey Ocyceros gingalensis
Image by: 1)
Lahiru_Prabudd_Fernando 2)
Rafael_Vila 3, 4) Gihan_Jayaweera
1) Female 2 - 4) Male
Genus Penelopides
Hornbill,_Luzon Penelopides manillae Found: Philippines
Image by: 1)
Mark A Harper 2)
Len_Worthington 3)
BS Thurner Hof
1) Female 2, 3) Male
Hornbill,_Mindanao Penelopides affinis Found: Philippines
Image by:
1)
Berstein49 3)
Blake Matheson 3)
Jovic_Ferrer
1) Female 2, 3) Male
Hornbill,_Samar Penelopides samarensis Found: Philippines
Image by: 1) Sergey_Yeliseev
Hornbill,_Visayan Penelopides panini Found: Philippines
Image by: 1)
Magalhães 2)
Nasser_Halaweh 3)
Steven_Wilson 4)
Tim Ellis - Chester Zoo
1, 2) Pair 3, 4) Male
Genus Rhabdotorrhinus
Hornbill,_Sulawesi Rhabdotorrhinus exarhatus Found: Indonesia
Image by: 1)
Vachovec1 2)
Chris
huh 3)
TR_Shankar_Raman
1) Pair
2, 3) Male
Hornbill,_Walden's Rhabdotorrhinus waldeni
Image by:
1)
Prof. Curio 2
, 3) Callen Bentley
Hornbill,_Wrinkled Rhabdotorrhinus corrugatus Found: southeast Asia
Image by: 1)
Linda_De_Volder 2)
Erik van Roekel 3)
Tony_Castro 4)
David_Cook - Borneo
1) Female 2 - 4) Male
Hornbill,_Writhed Rhabdotorrhinus leucocephalus Found, Philippines
Image by: 1)
John Gerrard Keulemans 2)
Francesco Veronesi 3)
Vanessa_David 3)
Patricia Casteren
1) Male on left 2) Female 3) Male
Genus Rhyticeros
These medium to large hornbills are found from Southeast Asia to the Solomons.
They have low, wreathed (wrinkled) casques and a mostly dull whitish horn-colored bill. They have mainly black plumage and most have a white tail. They have inflatable skin on the throat, which is blue for most males. The head and neck of the males are white or rufous.
Hornbill,_Blyth's Rhyticeros plicatus
Image by:
1)
Cifor 2)
yumievriwan 3)
Stephen Hanafin 4)
Jerry Oldenettel - Papua New Guinea
1, 2) Female
3) Male 4) Pair
Hornbill,_Knobbed Rhyticeros cassidix
Image by: 1)
WarriorWoman 2)
Mario Pineda - Los Angeles Zoo 3)
Lip Kee - North Sulawesi 4)
Asim_Bharwani
1) Pair 2) Female 3, 4) Male
Hornbill,_Narcondam Rhyticeros narcondami Found: Indian island of Narcondam
Image by:
1, 2)
Rohitjahnavi
1) Female 2) Male
Hornbill,_Plain-pouched Rhyticeros subruficollis
Image by: 1)
John Gerrard Keulemans
Hornbill,_Sumba Rhyticeros everetti
Image by: 1)
Stavenn
1) Female
Hornbill,_Wreathed Rhyticeros undulatus
Image by:
1)
Dick_Culbert 2)
Sandy Cole 3) Dick Daniels -
Sylvan Heights 4)
Jerry Oldenettel - Thailand
1, 2) Female 3, 4) Male
Genus Tockus
The hornbills in Lophoceros are close relatives and used to be include in genus tockus.
Hornbill,_Jackson's Tockus jacksoni
Image by:
1, 3) Charles_J_Sharp - Kenya
2) Steve Garvie 4)
Mathieu_Breitenstein - Kenya
1, 2) Female 3, 4) Male
Hornbill,_Monteiro's Tockus monteiri
Image by: 1)
Gregoire_Dubois 2)
Jeffrey Hotson - Namibia 3)
Nanosanchez
Hornbill,_von der Decken's Tockus deckeni
Image by
1)
Steven G Johnson - the London Zoo
2)
Dick Daniels -
Tampa's Lowry Park Zoo 3, 4) Nik_Borrow - Tanzania, Ethiopia
1, 2, 3) Female 4, 5, 6) Male
Red-billed Hornbills
The red-billed hornbill has been split into 5 species:
Damara red-billed hornbill,
northern red-billed hornbill,
southern red-billed hornbill,
Tanzanian red-billed hornbill, and
western red-billed hornbill. The red-hornbills have wings covered in circles of white feathers surrounded by black or dark brown feathers. These five species are listed next for ease of reference.
Hornbill,_Damara_Red-billed Tockus damarensis
Image by: 1)
Hans Hillewaert - Namibia 2)
Ron_Knight 3)
Graham_Duggan
Hornbill,_Northern_Red-billed Tockus erythrorhynchus Found: Africa
Image by: 1)
Alan_Hopkins - Tanzania
2, 3) Nik_Borrow - Ethiopia
2) Male
Hornbill,_Southern_Red-billed Tockus rufirostris
Image by: 1)
Dick Daniels - Kruger National Park, South Africa 2)
Arno_Meintjes 3)
Charles_J_Sharp - Kenya 4)
Tim_Ellis
Hornbill,_Tanzanian Red-billed Tockus ruahae
Image by:
1) Dick Daniels- Tanzaia 2)
Nevit Dilmen - Tanzania
3, 4)
Nik_Borrow - Tanzania
Hornbill,_Western Red-billed Tockus kempi
Image by:
1)
Allan Hopkins - Gambia 2) Thomas Brown 3)
Charles J Sharp - Gambia 4)
Nik_Borrow - Senegal
Yellow-billed Hornbills
The yellow-billed hornbill has been split into 2 species:
eastern yellow-billed hornbill and
southern yellow-billed hornbill. These species are listed next for ease of reference.
Hornbill,_Eastern_Yellow-billed Tockus flavirostris Found:
Africa
Image by: 1)
Eric - Discovery Cove, Orlando, Florida 2)
Greg Hume - Cincinnati Zoo 3)
Steve_Garvie 4)
Nik_Borrow - Ethiopia
Hornbill,_Southern Yellow-billed Tockus leucomelas Found:
Africa
Image by: 1)
Derek_Keats - South Africa 2)
Yathin_Krishnappa 3) Dick - South Africa 4)
Cristiano Crolle - Namibia
GROUND HORNBILLs
Order Bucerotiformes Family Bucorvidae
The family Bucorvidae has only two members, the Abyssinian ground-hornbill and the southern ground-hornbill. They are both found in Sub-Saharan Africa. Ground hornbills are large, with adults around a meter tall. Both species are ground-dwelling, unlike other hornbills, and feed on insects, snakes, other birds, amphibians and even tortoises. They only fly when they are alarmed.
The Abyssinian ground-hornbill in captivity lives 35 ot 40 years.
The larger southern species is possibly the slowest-breeding species and one of the longest-lived of all birds.They eat a wide variety of small vertebrates and invertebrates, including tortoises, lizards, snakes, birds, spiders, beetles, and caterpillars; they also take carrion, some fruits, seeds, and groundnuts.
Similar to the hornbills of family Bucerotidae, the ground-hornbills nest mainly in tree cavities, but also holes in rocks. These cavities are also partially sealed when the female is ready to lay eggs. A mixture of mud and vegetation is used accomplish the closure. Prior to construction, the male lines the cavity with dry leaves. The female usually lays 1 to 2 eggs with a three to five day separation between the eggs. The first born chick is usually 3 to 5 days older than its sibling; as result it is considerably larger and usually gets the majority of food so the younger chick dies. In essence, the second chick was an insurance policy. If the first born is defective, then chick number 2 gets lucky, otherwise life is hard.